:root {
  --green-950: #09251d;
  --green-900: #0f3b2e;
  --green-800: #15543f;
  --green-700: #106b4d;
  --green-500: #4caf72;
  --green-300: #9be0ac;
  --green-100: #e7f4e9;
  --cream: #f7f3e8;
  --sand: #efe6d2;
  --ink: #16251e;
  --muted: #5e6d63;
  --line: #dfe7df;
  --white: #ffffff;
  --shadow: 0 22px 70px rgba(9, 37, 29, .14);
  --radius-lg: 28px;
  --radius-md: 18px;
  --radius-sm: 12px;
  --container: 1180px;
}

* { box-sizing: border-box; }
html { scroll-behavior: smooth; }
body {
  margin: 0;
  font-family: ui-sans-serif, system-ui, -apple-system, BlinkMacSystemFont, "Segoe UI", sans-serif;
  color: var(--ink);
  background: var(--white);
  line-height: 1.55;
}
a { color: inherit; }
img { max-width: 100%; display: block; }
.container { width: min(calc(100% - 40px), var(--container)); margin: 0 auto; }
.narrow { max-width: 850px; }
.skip-link { position: absolute; left: -999px; top: 12px; z-index: 1000; background: var(--green-900); color: var(--white); padding: 10px 14px; border-radius: 8px; }
.skip-link:focus { left: 12px; }

.site-header {
  position: sticky;
  top: 0;
  z-index: 100;
  background: rgba(247, 243, 232, .88);
  backdrop-filter: blur(18px);
  border-bottom: 1px solid rgba(15, 59, 46, .08);
  transition: box-shadow .2s ease, background .2s ease;
}
.site-header.scrolled { box-shadow: 0 12px 38px rgba(9, 37, 29, .09); background: rgba(255,255,255,.92); }
.header-inner { display: flex; align-items: center; justify-content: space-between; min-height: 82px; gap: 24px; }
.brand { display: flex; align-items: center; gap: 12px; text-decoration: none; font-weight: 800; color: var(--green-900); }
.brand-mark { width: 156px; height: auto; }
.brand-text { display: none; }
.site-nav { display: flex; align-items: center; gap: 6px; }
.nav-link { text-decoration: none; font-size: 14px; font-weight: 700; color: var(--green-900); padding: 10px 13px; border-radius: 999px; transition: background .2s ease, color .2s ease; }
.nav-link:hover, .nav-link.active { background: var(--green-100); color: var(--green-700); }
.nav-toggle { display: none; background: transparent; border: 0; padding: 8px; cursor: pointer; }
.nav-toggle span { display: block; width: 24px; height: 2px; background: var(--green-900); margin: 5px 0; }

.btn { display: inline-flex; align-items: center; justify-content: center; gap: 8px; text-decoration: none; border-radius: 999px; font-weight: 800; padding: 14px 20px; border: 1px solid transparent; transition: transform .2s ease, box-shadow .2s ease, background .2s ease; }
.btn:hover { transform: translateY(-1px); }
.btn-primary { background: var(--green-900); color: var(--white); box-shadow: 0 14px 34px rgba(15, 59, 46, .22); }
.btn-secondary { background: var(--white); color: var(--green-900); border-color: rgba(15,59,46,.16); }
.btn-outline { border-color: rgba(15,59,46,.25); color: var(--green-900); background: transparent; }
.btn-light { background: var(--cream); color: var(--green-900); }
.btn-small { padding: 10px 14px; font-size: 14px; }

.hero { background: radial-gradient(circle at 82% 8%, rgba(155,224,172,.45), transparent 28%), linear-gradient(135deg, var(--cream) 0%, #fdfbf5 52%, #e9f4eb 100%); overflow: hidden; }
.hero-grid { min-height: 720px; display: grid; grid-template-columns: 1.06fr .94fr; gap: 56px; align-items: center; padding: 82px 0; }
.hero-copy h1, .page-hero h1 { margin: 0 0 24px; font-size: clamp(50px, 7vw, 92px); line-height: .92; letter-spacing: -4px; color: var(--green-950); }
.hero-subhead, .page-hero p, .lead { font-size: clamp(18px, 2vw, 22px); color: #415247; }
.hero-actions { display: flex; gap: 12px; flex-wrap: wrap; margin-top: 34px; }
.eyebrow { margin: 0 0 14px; color: var(--green-700); font-size: 13px; font-weight: 900; letter-spacing: .14em; text-transform: uppercase; }
.hero-visual { position: relative; min-height: 500px; display: flex; align-items: center; justify-content: center; }
.visual-card { background: rgba(255,255,255,.78); border: 1px solid rgba(15,59,46,.1); border-radius: 34px; box-shadow: var(--shadow); padding: 16px; }
.visual-card-large { transform: rotate(-1deg); }
.status-card { position: absolute; width: 190px; background: var(--white); border: 1px solid rgba(15,59,46,.1); border-radius: 18px; padding: 16px; box-shadow: 0 18px 45px rgba(9,37,29,.13); }
.status-card strong { display: block; font-size: 18px; color: var(--green-900); }
.status-label { display: block; color: var(--muted); font-size: 13px; font-weight: 800; text-transform: uppercase; letter-spacing: .08em; }
.status-one { left: -2px; top: 72px; }
.status-two { right: -8px; bottom: 76px; }

.proof-strip { background: var(--green-950); color: var(--white); }
.proof-grid { display: grid; grid-template-columns: repeat(4, 1fr); }
.proof-grid div { padding: 26px 22px; border-left: 1px solid rgba(255,255,255,.12); }
.proof-grid div:last-child { border-right: 1px solid rgba(255,255,255,.12); }
.proof-grid span { display: block; color: rgba(255,255,255,.62); font-size: 12px; letter-spacing: .12em; text-transform: uppercase; font-weight: 800; }
.proof-grid strong { display: block; margin-top: 5px; font-size: 18px; }

.section { padding: 104px 0; }
.section-tight { padding-top: 40px; }
.section-light { background: #f7f8f2; }
.section-dark { background: radial-gradient(circle at 15% 0%, rgba(76,175,114,.25), transparent 30%), var(--green-950); color: var(--white); }
.section-cta { padding: 80px 0; background: linear-gradient(135deg, var(--green-800), var(--green-950)); color: var(--white); }
.split-grid { display: grid; grid-template-columns: .84fr 1.16fr; gap: 72px; }
.split-grid.reverse { grid-template-columns: 1.08fr .92fr; }
.align-center { align-items: center; }
.section h2, .page-hero h1 { letter-spacing: -2px; }
.section h2 { margin: 0; font-size: clamp(34px, 4vw, 56px); line-height: 1.02; color: var(--green-950); }
.section-dark h2, .section-cta h2, .light-heading h2 { color: var(--white); }
.section p { color: var(--muted); }
.section-dark p, .section-cta p, .light-heading p { color: rgba(255,255,255,.78); }
.rich-text p:first-child { margin-top: 0; }
.rich-text p { font-size: 18px; }
.centered { text-align: center; margin-bottom: 44px; }
.section-heading { max-width: 820px; margin-bottom: 44px; }
.section-heading.centered { margin-left: auto; margin-right: auto; }
.section-heading p { font-size: 18px; }
.card-grid { display: grid; gap: 22px; }
.card-grid.three { grid-template-columns: repeat(3, 1fr); }
.card-grid.four { grid-template-columns: repeat(4, 1fr); }
.card { background: var(--white); border: 1px solid var(--line); border-radius: var(--radius-lg); padding: 28px; box-shadow: 0 14px 40px rgba(9,37,29,.06); }
.card h3 { margin: 18px 0 10px; color: var(--green-950); font-size: 23px; line-height: 1.1; }
.icon-pill { display: inline-flex; align-items: center; justify-content: center; width: 42px; height: 42px; border-radius: 999px; background: var(--green-100); color: var(--green-700); font-weight: 900; }
.panel-glow { padding: 42px; border-radius: var(--radius-lg); background: rgba(255,255,255,.08); border: 1px solid rgba(255,255,255,.14); }
.panel-kicker { margin-top: 0; font-size: 13px; font-weight: 900; letter-spacing: .14em; text-transform: uppercase; color: var(--green-300) !important; }
.value-grid { display: grid; grid-template-columns: repeat(2, 1fr); gap: 16px; }
.value-grid div, .partner-models div { padding: 24px; border-radius: var(--radius-md); background: rgba(255,255,255,.08); border: 1px solid rgba(255,255,255,.13); }
.value-grid strong, .partner-models strong { display: block; font-size: 22px; color: var(--white); }
.value-grid span, .partner-models span { color: rgba(255,255,255,.73); }
.timeline-preview { display: grid; gap: 14px; position: relative; }
.timeline-preview div { position: relative; padding: 24px 28px; background: #f7f8f2; border: 1px solid var(--line); border-radius: var(--radius-md); font-weight: 900; color: var(--green-900); }
.timeline-preview div::before { content: ""; position: absolute; left: -14px; top: 50%; width: 14px; height: 2px; background: var(--green-500); }
.cta-card { display: flex; align-items: center; justify-content: space-between; gap: 32px; padding: 44px; border-radius: 34px; background: rgba(255,255,255,.1); border: 1px solid rgba(255,255,255,.16); }
.cta-card h2 { margin: 0 0 14px; font-size: clamp(32px, 4vw, 54px); line-height: 1; }
.cta-card p { margin-bottom: 0; max-width: 740px; }

.page-hero { padding: 128px 0 88px; background: radial-gradient(circle at 80% 8%, rgba(155,224,172,.45), transparent 28%), linear-gradient(135deg, var(--cream), #fffdf7 70%); }
.page-hero h1 { max-width: 900px; }
.page-hero p { max-width: 820px; }
.technology-grid { display: grid; grid-template-columns: .9fr 1.1fr; gap: 62px; align-items: center; }
.technology-diagram { padding: 12px; }
.process-grid { display: grid; grid-template-columns: repeat(4, 1fr); gap: 18px; }
.process-step { padding: 26px; border-radius: var(--radius-lg); background: #fbfcf8; border: 1px solid var(--line); }
.process-step span { display: inline-flex; align-items: center; justify-content: center; width: 38px; height: 38px; border-radius: 50%; background: var(--green-900); color: var(--white); font-weight: 900; }
.process-step h3 { font-size: 21px; line-height: 1.15; }
.dark-card { background: rgba(255,255,255,.08); border-color: rgba(255,255,255,.14); box-shadow: none; }
.dark-card h3 { color: var(--white); }
.dark-card p { color: rgba(255,255,255,.72); }
.application-list { display: grid; gap: 18px; }
.application-list article { padding: 26px; border-radius: var(--radius-md); border: 1px solid var(--line); background: var(--white); }
.application-list h3 { margin: 0 0 8px; color: var(--green-900); }

.pipeline-stages { display: grid; grid-template-columns: repeat(5, 1fr); gap: 14px; counter-reset: stage; }
.pipeline-stages article { position: relative; padding: 24px; background: var(--white); border: 1px solid var(--line); border-radius: var(--radius-md); min-height: 260px; }
.pipeline-stages span { display: block; color: var(--green-500); font-weight: 900; letter-spacing: .1em; }
.pipeline-stages h3 { font-size: 21px; line-height: 1.1; color: var(--green-950); }
.program-list { display: grid; gap: 22px; }
.program-card { display: grid; grid-template-columns: 1.1fr .9fr; gap: 28px; padding: 34px; border: 1px solid var(--line); border-radius: var(--radius-lg); background: var(--white); box-shadow: 0 14px 40px rgba(9,37,29,.06); }
.featured-program { border-color: rgba(16,107,77,.32); background: linear-gradient(135deg, #ffffff, #f1faef); }
.program-label { margin: 0 0 10px; color: var(--green-700) !important; font-size: 12px; font-weight: 900; letter-spacing: .12em; text-transform: uppercase; }
.program-card h3 { margin: 0 0 12px; font-size: 30px; line-height: 1.05; color: var(--green-950); }
.program-details { display: grid; gap: 12px; margin: 0; }
.program-details div { padding: 15px 18px; border-radius: var(--radius-sm); background: #f7f8f2; }
.program-details dt { font-size: 12px; font-weight: 900; letter-spacing: .1em; text-transform: uppercase; color: var(--muted); }
.program-details dd { margin: 3px 0 0; color: var(--green-950); font-weight: 800; }
.partner-models { display: grid; gap: 14px; }

.team-grid { display: grid; grid-template-columns: repeat(3, 1fr); gap: 22px; }
.team-card { padding: 30px; border: 1px solid var(--line); border-radius: var(--radius-lg); background: var(--white); box-shadow: 0 14px 38px rgba(9,37,29,.06); }
.avatar { width: 78px; height: 78px; border-radius: 24px; display: flex; align-items: center; justify-content: center; background: linear-gradient(135deg, var(--green-100), var(--green-300)); color: var(--green-900); font-size: 26px; font-weight: 900; margin-bottom: 22px; }
.team-card h3 { margin: 0; font-size: 25px; color: var(--green-950); }
.team-card .role { margin: 4px 0 14px; color: var(--green-700); font-weight: 900; }
.credibility-grid { display: grid; grid-template-columns: 1fr .8fr; gap: 60px; align-items: center; }
.cred-card { padding: 36px; border-radius: var(--radius-lg); background: rgba(255,255,255,.08); border: 1px solid rgba(255,255,255,.14); }
.cred-card strong { display: block; font-size: 30px; margin-bottom: 8px; }
.cred-card span { color: rgba(255,255,255,.75); }

.news-feature { display: flex; justify-content: space-between; align-items: center; gap: 28px; padding: 34px; background: var(--white); border-radius: var(--radius-lg); border: 1px solid var(--line); box-shadow: 0 14px 38px rgba(9,37,29,.06); }
.news-feature h2 { margin-bottom: 12px; }
.news-grid { display: grid; grid-template-columns: repeat(2, 1fr); gap: 22px; }
.news-card { padding: 30px; border: 1px solid var(--line); border-radius: var(--radius-lg); background: var(--white); box-shadow: 0 14px 38px rgba(9,37,29,.06); }
.news-date { display: block; color: var(--green-700); font-size: 12px; font-weight: 900; letter-spacing: .12em; text-transform: uppercase; margin-bottom: 14px; }
.news-card h3 { margin: 0 0 12px; color: var(--green-950); font-size: 26px; line-height: 1.08; }
.news-card a { color: var(--green-700); font-weight: 900; text-decoration: none; }
.news-card a:hover { text-decoration: underline; }
.article-layout { display: grid; grid-template-columns: minmax(0, 1fr) 330px; gap: 58px; align-items: start; }
.article-body { font-size: 19px; }
.article-body h2 { font-size: 32px; margin-top: 44px; }
.article-body blockquote { margin: 36px 0; padding: 28px; border-left: 5px solid var(--green-500); background: #f7f8f2; border-radius: 0 var(--radius-md) var(--radius-md) 0; color: var(--green-950); font-weight: 700; }
.article-note { position: sticky; top: 110px; padding: 24px; border-radius: var(--radius-md); background: var(--green-100); border: 1px solid rgba(16,107,77,.18); }
.article-note h2 { font-size: 22px; margin-top: 0; }
.article-note li { margin: 10px 0; }

.site-footer { background: var(--green-950); color: var(--white); padding: 66px 0 26px; }
.footer-grid { display: grid; grid-template-columns: 1.1fr .9fr .7fr; gap: 60px; }
.footer-brand { display: inline-flex; align-items: center; gap: 12px; color: var(--white); text-decoration: none; font-size: 20px; font-weight: 900; }
.footer-mark { width: 54px; height: auto; filter: brightness(0) invert(1); opacity: .95; }
.footer-copy, .site-footer p { color: rgba(255,255,255,.72); }
.footer-heading { font-size: 15px; text-transform: uppercase; letter-spacing: .12em; color: var(--green-300); }
.footer-link { color: var(--white); font-weight: 800; }
.footer-bottom { display: flex; justify-content: space-between; gap: 20px; border-top: 1px solid rgba(255,255,255,.12); margin-top: 52px; padding-top: 22px; color: rgba(255,255,255,.6); font-size: 14px; }
.fine-print { font-size: 13px; }

@media (max-width: 1040px) {
  .hero-grid, .split-grid, .split-grid.reverse, .technology-grid, .credibility-grid { grid-template-columns: 1fr; }
  .hero-grid { min-height: auto; }
  .hero-visual { min-height: 420px; }
  .card-grid.four, .process-grid, .pipeline-stages { grid-template-columns: repeat(2, 1fr); }
  .team-grid { grid-template-columns: repeat(2, 1fr); }
  .footer-grid { grid-template-columns: 1fr 1fr; }
}

@media (max-width: 820px) {
  .header-inner { min-height: 72px; }
  .nav-toggle { display: block; }
  .site-nav { position: fixed; inset: 72px 20px auto 20px; display: none; flex-direction: column; align-items: stretch; background: var(--white); padding: 18px; border-radius: 22px; box-shadow: var(--shadow); border: 1px solid var(--line); }
  .site-nav.open { display: flex; }
  .nav-link, .btn-small { width: 100%; justify-content: flex-start; }
  .brand-mark { width: 142px; }
  .hero-grid { padding: 56px 0; gap: 30px; }
  .hero-copy h1, .page-hero h1 { letter-spacing: -2px; }
  .proof-grid { grid-template-columns: repeat(2, 1fr); }
  .card-grid.three, .card-grid.four, .process-grid, .pipeline-stages, .team-grid, .news-grid { grid-template-columns: 1fr; }
  .program-card, .article-layout { grid-template-columns: 1fr; }
  .news-feature, .cta-card, .footer-bottom { flex-direction: column; align-items: flex-start; }
  .footer-grid { grid-template-columns: 1fr; gap: 28px; }
  .section { padding: 76px 0; }
  .page-hero { padding: 88px 0 66px; }
  .hero-visual { min-height: 360px; }
  .status-card { position: static; margin: 12px; width: auto; }
  .hero-visual { flex-direction: column; }
}

@media (max-width: 520px) {
  .container { width: min(calc(100% - 28px), var(--container)); }
  .hero-actions { flex-direction: column; align-items: stretch; }
  .btn { width: 100%; }
  .proof-grid { grid-template-columns: 1fr; }
  .value-grid { grid-template-columns: 1fr; }
  .cta-card { padding: 28px; }
  .card, .team-card, .program-card, .news-card { padding: 24px; }
}

/* Supplied Upstream Biotechnology wordmark integration */
.site-header {
  background: rgba(0, 0, 0, .96);
  border-bottom: 1px solid rgba(255, 255, 255, .10);
}
.site-header.scrolled {
  background: rgba(0, 0, 0, .98);
  box-shadow: 0 12px 38px rgba(0, 0, 0, .20);
}
.header-inner { min-height: 92px; }
.brand { color: var(--white); }
.brand-mark {
  width: 206px;
  height: auto;
}
.nav-link {
  color: rgba(255, 255, 255, .84);
}
.nav-link:hover,
.nav-link.active {
  background: rgba(76, 175, 114, .18);
  color: var(--white);
}
.site-header .btn-outline {
  border-color: rgba(255, 255, 255, .34);
  color: var(--white);
  background: rgba(255, 255, 255, .04);
}
.site-header .btn-outline:hover {
  background: rgba(255, 255, 255, .10);
  box-shadow: 0 12px 28px rgba(0, 0, 0, .18);
}
.nav-toggle span { background: var(--white); }
.site-footer { background: #000; }
.footer-brand span { display: none; }
.footer-mark {
  width: 206px;
  height: auto;
  filter: none;
  opacity: 1;
}

@media (max-width: 820px) {
  .header-inner { min-height: 82px; }
  .site-nav {
    inset: 82px 20px auto 20px;
    background: #000;
    border-color: rgba(255, 255, 255, .14);
  }
  .brand-mark { width: 184px; }
}

@media (max-width: 520px) {
  .brand-mark { width: 170px; }
  .footer-mark { width: 184px; }
}


/* Website refinement update */
.product-pipeline-section { align-items: center; }
.pipeline-figure { margin: 0; padding: 16px; background: #ffffff; border: 1px solid var(--line); border-radius: 28px; box-shadow: 0 18px 42px rgba(9,37,29,.08); }
.pipeline-figure img { width: 100%; height: auto; border-radius: 18px; }
.pipeline-figure figcaption { margin-top: 12px; font-size: 14px; color: var(--muted); }
.pipeline-visual-wrap { margin-top: 10px; margin-bottom: 34px; }
.pipeline-figure-wide { max-width: 1120px; }
.leadership-grid { display: grid; grid-template-columns: repeat(2, 1fr); gap: 22px; }
.science-grid { display: grid; grid-template-columns: repeat(3, 1fr); gap: 22px; }
.team-card { height: 100%; }
.team-card-leadership { display: grid; grid-template-columns: 150px 1fr; gap: 24px; align-items: center; }
.team-card-science { display: block; }
.portrait-shell { display: flex; align-items: center; justify-content: center; overflow: hidden; background: radial-gradient(circle at top, rgba(155,224,172,.28), rgba(247,243,232,.95)); border: 1px solid rgba(15,59,46,.08); box-shadow: inset 0 1px 0 rgba(255,255,255,.65); }
.portrait-shell img { width: 100%; height: 100%; object-fit: cover; }
.portrait-large { width: 150px; height: 150px; border-radius: 50%; margin: 0 auto; }
.portrait-small { width: 108px; height: 108px; border-radius: 50%; margin-bottom: 18px; }
.team-card-science h3 { font-size: 22px; }
.scientific-team-section .section-heading { margin-bottom: 30px; }
@media (max-width: 1040px) {
  .leadership-grid, .science-grid { grid-template-columns: repeat(2, 1fr); }
}
@media (max-width: 820px) {
  .leadership-grid, .science-grid { grid-template-columns: 1fr; }
  .team-card-leadership { grid-template-columns: 1fr; text-align: center; }
  .portrait-small { margin-left: auto; margin-right: auto; }
}


/* Leadership roster update */
@media (min-width: 1041px) {
  .leadership-grid { grid-template-columns: repeat(2, 1fr); }
  .leadership-grid .team-card-leadership:last-child { grid-column: 1 / -1; }
  .leadership-grid .team-card-leadership:last-child .portrait-large { width: 138px; height: 138px; }
}


/* v4 refined approach and founder bios */
.approach-localization {
  margin-top: 34px;
  display: grid;
  grid-template-columns: .78fr 1.22fr;
  gap: 38px;
  align-items: center;
  padding: 34px;
  border-radius: 30px;
  background: linear-gradient(135deg, #f7f8f2, #ffffff);
  border: 1px solid rgba(16,107,77,.18);
  box-shadow: 0 16px 44px rgba(9,37,29,.06);
}
.approach-localization h3 {
  margin: 0;
  font-size: clamp(30px, 4vw, 48px);
  line-height: 1.02;
  letter-spacing: -1.5px;
  color: var(--green-950);
}
.approach-localization .rich-text p { font-size: 17px; }
.cofounder-grid { display: grid; grid-template-columns: repeat(2, 1fr); gap: 24px; }
.operations-grid { display: grid; grid-template-columns: repeat(3, 1fr); gap: 22px; }
.team-card-cofounder { display: grid; grid-template-columns: 160px 1fr; gap: 26px; align-items: start; padding: 34px; border-color: rgba(16,107,77,.24); background: linear-gradient(135deg, #ffffff, #f7fbf4); }
.team-card-cofounder h3 { font-size: 30px; }
.team-card-cofounder p { font-size: 16px; }
.portrait-founder { width: 160px; height: 160px; border-radius: 50%; margin: 0 auto; }
.operations-grid .team-card { box-shadow: 0 12px 34px rgba(9,37,29,.055); }
.team-card-operations { display: block; }
.portrait-medium { width: 132px; height: 132px; border-radius: 50%; margin-bottom: 18px; }
.leadership-support-section .section-heading { margin-bottom: 30px; }
@media (max-width: 1120px) {
  .cofounder-grid, .operations-grid { grid-template-columns: 1fr; }
  .team-card-cofounder { grid-template-columns: 150px 1fr; }
  .portrait-founder { width: 150px; height: 150px; }
}
@media (max-width: 820px) {
  .team-card-cofounder { grid-template-columns: 1fr; text-align: center; }
  .portrait-founder, .portrait-medium { margin-left: auto; margin-right: auto; }
}


/* v5 homepage approach follow-on */
.approach-followon {
  margin-top: 76px;
  padding-top: 66px;
  border-top: 1px solid var(--line);
}
.approach-followon h2 { color: var(--green-950); }


/* Interactive RNA SwitchBlade graphic prototype */
.interactive-visual-card, .interactive-tech-diagram { padding-bottom: 12px; }
.interactive-rna-graphic { position: relative; width: min(100%, 720px); cursor: pointer; }
.rnaswitch-svg { display: block; width: 100%; height: auto; cursor: pointer; }
.hover-hint { margin: 14px 6px 2px; font-size: 13px; line-height: 1.45; color: var(--green-700) !important; font-weight: 700; }
.interactive-rna-graphic::after { content: ""; position: absolute; inset: 0; border-radius: 26px; pointer-events: none; box-shadow: inset 0 0 0 1px rgba(15,59,46,.04); }
.interactive-rna-graphic .logic-green-bar,
.interactive-rna-graphic .disease-pressure-graphics,
.interactive-rna-graphic .disease-wave,
.interactive-rna-graphic .disease-dots,
.interactive-rna-graphic .stress-pill {
  transition: opacity .22s ease, fill .22s ease, stroke .22s ease, filter .22s ease;
}
.interactive-rna-graphic .logic-green-bar {
  fill: #9be0ac;
}
.interactive-rna-graphic .disease-pressure-graphics {
  opacity: .62;
}
.interactive-rna-graphic.is-right-active .logic-green-bar {
  fill: #57ff88;
  filter: url(#barGlowProto);
}
.interactive-rna-graphic.is-right-active .disease-pressure-graphics {
  animation: diseasePulse 1.5s ease-in-out infinite;
  filter: url(#diseaseGlowProto);
}
@keyframes diseasePulse {
  0%, 100% { opacity: .35; }
  50% { opacity: 1; }
}
@media (hover: none) {
  .interactive-rna-graphic .logic-green-bar {
    fill: #b0e8bc;
  }
  .interactive-rna-graphic .disease-pressure-graphics {
    opacity: .78;
  }
}

/* Team bio portrait hover treatment triggered by bio text */
.team-card { overflow: visible; }
.portrait-shell {
  position: relative;
  z-index: 1;
  transition: transform .22s ease, box-shadow .22s ease, border-color .22s ease;
  transform-origin: center center;
}
.bio-hover-trigger {
  cursor: pointer;
}
.portrait-shell.bio-active {
  transform: scale(1.5);
  z-index: 8;
  box-shadow: 0 26px 70px rgba(9,37,29,.30), 0 8px 20px rgba(9,37,29,.18);
  border-color: rgba(16,107,77,.28);
}
@media (hover: none) {
  .portrait-shell.bio-active {
    transform: none;
    box-shadow: inset 0 1px 0 rgba(255,255,255,.65);
  }
}

/* Technology page title size alignment */
.technology-hero-title {
  font-size: clamp(34px, 4vw, 56px) !important;
  line-height: 1.02 !important;
}

/* News detail pages */
.article-source-link { overflow-wrap: anywhere; word-break: break-word; }
.article-note a { color: var(--green-700); font-weight: 800; }


/* Technology explainer animation */
.technology-explainer-section {
  background: linear-gradient(180deg, #f7fbf4 0%, #ffffff 100%);
}
.technology-video-card {
  max-width: 1080px;
  margin: 0 auto;
  padding: 18px;
  border: 1px solid var(--line);
  border-radius: 32px;
  background: var(--white);
  box-shadow: 0 22px 60px rgba(9,37,29,.10);
}
.technology-explainer-video {
  display: block;
  width: 100%;
  aspect-ratio: 16 / 9;
  object-fit: cover;
  border-radius: 22px;
  background: #f6efdb;
}
.technology-video-card figcaption {
  padding: 16px 8px 4px;
  color: #526258;
  font-size: 15px;
  line-height: 1.55;
}
@media (max-width: 820px) {
  .technology-video-card { padding: 10px; border-radius: 22px; }
  .technology-explainer-video { border-radius: 14px; }
  .technology-video-card figcaption { padding: 12px 6px 2px; }
}


/* Publications and science section */
.science-section { border-top: 1px solid var(--line); }
.publication-feature-grid { display: grid; grid-template-columns: repeat(3, minmax(0, 1fr)); gap: 22px; }
.publication-card { display: flex; flex-direction: column; min-height: 330px; padding: 30px; border: 1px solid var(--line); border-radius: var(--radius-lg); background: var(--white); box-shadow: 0 14px 38px rgba(9,37,29,.06); }
.publication-card-featured { background: linear-gradient(145deg, #ffffff, #f3f8ef); }
.publication-year { display: block; margin-bottom: 16px; color: var(--green-700); font-size: 12px; font-weight: 900; letter-spacing: .12em; text-transform: uppercase; }
.publication-card h3 { margin: 0 0 18px; color: var(--green-950); font-size: 24px; line-height: 1.12; }
.publication-card p { margin: 0 0 24px; color: var(--muted); }
.publication-paper-link { margin-top: auto; color: var(--green-700); font-weight: 900; text-decoration: none; }
.publication-paper-link:hover { text-decoration: underline; }
.publication-more { display: flex; justify-content: center; margin-top: 34px; }
.publication-list-heading { max-width: 820px; margin-bottom: 44px; }
.publication-list-heading h2 { margin: 0; font-size: clamp(34px, 4vw, 56px); line-height: 1.02; color: var(--green-950); letter-spacing: -2px; }
.publication-list { display: grid; gap: 16px; }
.publication-list-item { display: grid; grid-template-columns: minmax(0, 1fr) auto; gap: 30px; align-items: center; padding: 26px 28px; border: 1px solid var(--line); border-radius: var(--radius-md); background: var(--white); box-shadow: 0 10px 28px rgba(9,37,29,.045); }
.publication-list-item h2 { margin: 0 0 8px; color: var(--green-950); font-size: 23px; line-height: 1.18; letter-spacing: -.5px; }
.publication-list-item p { margin: 0; color: var(--muted); }
.publication-list-button { white-space: nowrap; }
@media (max-width: 820px) {
  .publication-feature-grid { grid-template-columns: 1fr; }
  .publication-card { min-height: auto; }
  .publication-list-item { grid-template-columns: 1fr; gap: 18px; }
  .publication-list-button { width: 100%; }
}

/* July 10 reconstruction: publications Load more */
.more-publications {
  display: grid;
  gap: 16px;
}
.more-publications[hidden] {
  display: none;
}
.publication-load-more {
  display: flex;
  justify-content: center;
  margin-top: 34px;
}
.publication-load-more-button {
  min-width: 150px;
}



/* v34 publishing-readiness corrections */
body.nav-open { overflow: hidden; }
.page-hero h1 { max-width: 100%; overflow-wrap: anywhere; word-break: normal; }
img, video, svg { max-width: 100%; }
@media (max-width: 520px) {
  .page-hero h1 {
    font-size: clamp(36px, 10.5vw, 48px);
    line-height: .98;
    letter-spacing: -1.5px;
  }
}